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ARA-290 operates through three complementary mechanisms identified across pre-clinical and clinical research: 1

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Depending on the substrate in question, these steps can be performed with or without participation of a carrier lipid and, in some cases, are stimulated by the bacterial second messenger bis-(35)-cyclic dimeric guanosine monophosphate (c-di-GMP 49 )

J Infect Dis 185: 405407