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I found the reference to stiffness and muscle pain in undertreated B12 deficiency....it's in the pinned posts to the right on the home screen (sorry, foggy so can't remember precisely which one - but it's there if you want to have a search)

Oxidative stress and accelerated aging are the clearest targets, since glutathione is the front-line molecule that neutralizes free radicals before they damage mitochondria and DNA

10.1016/j.annonc.2020.02.011 101 LiuQ.WangS.PeiG.YangY.HuangY

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Sperm capacitation and acrosome reaction in mammalian sperm

Even the review paper summarizing this study noted the results are not overly informative and reliable due to the lack of objective measures